Ph.D., University of Texas at Austin, 1973.
380 Life Sciences Building
Department of Biological Sciences
Louisiana State University
Baton Rouge, Louisiana 70803 USA
Phone: 225-388-8551 and 225-388-8562
Mycology at LSU
Errata for Alexopoulos
Alexopoulos Text-Third Printing
Items for Alexopoulos Text
Fungal systematics and evolution; life histories; arthropod-dispersal of fungi
My interest in the phylogeny of fungi focuses on the way
in which arthropod dispersal has evolved. The problems associated
with spore dispersal of fungi to particular limited substrates
such as dung, have led to a broader investigation of spore
dispersal by arthropods. Fungi associated with mites phoretic
on beetles are especially important to provide new clues to
the evolution of entomogenous fungi; studies of phylogeny
of these groups is being emphasized. Enlargement of the collections
of unique Gulf Coastal Plain fungi, particularly wood-rotting
Basidiomycetes and fungi associated with arthropods is under
way. Fungi are studied using a variety of techniques including
field and cultural studies, electron microscopy, and nucleic
Department of Biological Sciences
Louisiana State University
Baton Rouge, LA 70803
B.S. 1961 Biology, University of Southwestern Louisiana, Lafayette
M.S. 1963 Biology, University of Alabama, Tuscaloosa. Fishes
of the Cahaba River Drainage System of Alabama
Ph.D. 1973 Botany, University of Texas at Austin. A Developmental
and Taxonomic Study of Protophysarum phloiogenum.
1962-1963 Graduate Teaching Fellow, University of Alabama
1962-1963 Graduate Teaching Assistant, University of Texas
1963-1968 Graduate Research Assistant, University of Texas
1970-1972 Graduate Research Assistant, University of Texas
1972-1974 Electron Microscopist, University of Florida
1974-1975 Interim Assistant in Botany, University of Florida
1975 (Summer) Faculty, Mountain Lake Biological Station, University of
1975-1981 Assistant Professor of Biology, Hope College, Holland,
Michigan (tenure granted, 1981)
1981-1985 Assistant Professor, Department of Botany, Louisiana State
University, Baton Rouge
1985-1988 Associate Professor with tenure, Department of Botany,
Louisiana State University
1988-1997 Professor, Department of Botany (then Plant Biology, now
Biological Sciences), Louisiana State University, Baton Rouge
1997- Boyd Professor, Department of Biological Sciences), Louisiana
State University, Baton Rouge
Acting Associate Dean, College of Basic Science,
Louisiana State University (7 months, 1989-1990) Acting
Chair, Department of Botany, Louisiana State University
(periodically 1989- ) Program Director, National
Science Foundation (1997-1998). Additional experiences listed
under "Professional Service."
OTHER PROFESSIONAL EXPERIENCE
Research Associate, Brown and Root-Northrup (NASA, Houston)
(1970, 4 months)
Societies: Mycological Society of America (Member, Committee on Research
Grants and Publications, 1976-1979; Chairman, 1977-1979; Councilor, 1981-1982;
Mycologia Memoirs Editorial Board, 1983-1990; Secretary, 1986-1988; Committee
on Culture Collections, 1986-1988; Foray Committee, 1986-1988; Committee for
Long Range Planning, 1989-1990, 1999-2000; Committee on the By-Laws, 1997-1999);
Vice-President, 1990-1991; President-Elect, 1991-1992; President, 1992-1993;
Past-President, 1993-1995) International Mycological Association (Member,
Executive Committee, 1990-1994; Secretary-General, 1994-1998; President, 1998-2002)
British Mycological Society (Member) Linnean Society of London
(Fellow) Botanical Society of America (Member; Secretary, Microbiological
Section, 1979-1982; Section Program Chairman, 1980-1982) AAAS (Fellow)
Willi Hennig Society (Associate Member) Society for Invertebrate
Pathology (Member) International Symbiosis Society (Member) Sigma
Xi (Member; Treasurer, Local Chapter, 1977-1979; Member, Awards Committee,
Other: Molecular Phylogenetics and Evolution (Member, Editorial
Board, 1992- ) Mycologia (Associate Editor, 1995-1997) Journal
of African Mycology and Biotechnology (Member, Editorial Board, 1998-
) Systematics and Geography of Plants (Member, Editorial Board,
1999- ) XVI International Botanical Congress (Member, Steering Committee,
1995-1999) International Union of Microbiological Societies (Mycology
Section), Member, Congress Programme Advisory Board (1997-1998) International
Association of Botanical and Mycological Societies, Member, ad hoc Committee
to assess invitations to host the XVII IBC National Science Foundation
(Panelist: Biotic Surveys and Inventories; Systematics; Infrastructure;
PEET White Paper on Monographs Committee; Systematics and Population Biology
Committee of Visitors; Program Director, Systematics Program, 1997-1998) III
Latin American Mycological Congress (Member, International Advisory Committee,
Miami University, Reviewer, Department of Botany, 1996
University of Gent, Reader, PhD Thesis of A. DeKesel, 1997
Harvard University, Member, PhD Qualifying Exam Committee of Debbie Miller,
Uppsala University, Advisor to the Search Committee, Systematic Botany position,
University of Alberta, Reader, PhD Thesis of Sarah Hambleton, 1998
University of the OFS, Bloemfontein, RSA, Reader, PhD Thesis of R. C. Witthuhn,
University of British Columbia, Reviewer, Department of Botany, 1999
HONORS AND AWARDS
1962-1963 University Teaching
Fellowship, University of Alabama, Tuscaloosa
1963 summer Traineeship, NIH Environmental Training
Program, Tulane University, New Orleans, Louisiana
1972 summer Traineeship, NSF Environmental and
Systematic Biology Program, University of Texas, Austin
Prize, Mycological Society of America
Fellow, British Mycological Society
Undergraduate: Botany (Majors' and Non-majors' sequences), General
Biology, Cytology, Plant Morphology, General Zoology, Microbiology,
Scanning Electron Microscopy Techniques, Economic Plants.
Graduate: Mycology, Advanced Mycology
INVITED SYMPOSIUM PAPERS
Resistant structures in the Myxomycetes. Second International Fungal Spore
Symposium, Provo, Utah, July 1974 (with H. C. Aldrich).
Identification of airborne myxomycete spores. AIBS, Athens, Georgia, August
Fungi of the Sonoran Desert. MSA, Milestones in Mycology Program, Bloomington,
Indiana, August 1981 (with R. L. Gilbertson) (Abstract).
Historical studies on fungus-insect relationships: an introduction to phylogenetic
systematics. Fungus-Insect Relationships. ESA, Eastern Branch and Cornell
University, Syracuse and Ithaca, N.Y., October 1981 (with Q. Wheeler).
Myxomycete life cycles and arthropod spore dispersers. Fungus-Insect Relationships.
ESA, Eastern Branch and Cornell University, Syracuse and Ithaca, N.Y., October
Spore dispersal in soybean pathogenic fungi. Louisiana Society of Electron
Microscopists, Keynote Address, Thibodaux, Louisiana, November 1983.
Host and geographical distribution of Gulf Coast wood-rotting fungi. MSA,
Gainesville, Florida, August 1985 (with R. L. Gilbertson) (Abstract).
Fungal interfaces: ectoparasites of arthropods. MSA, Amherst, Massachusetts,
August 1986 (Abstract).
Influence of arthropod dispersal on the evolution of fungal life cycles.
Fungal-insect interactions: a mini-symposium, University of Oregon, Eugene,
Oregon. April-May 1987.
Origins of invertebrate pathogens: Fungi. Plenary Session, Society for Invertebrate
Pathology, College Park, Maryland, August 1989 (with D. Malloch).
Taxonomy and phylogeny of Pyxidiophora: Molecular evidence. Taxonomy
and Biology of the Ophiostomatales Symposium, Bad Windsheim, West Germany,
August 1990 (with J.W. Spatafora, D. Malloch, and J.W. Taylor) (Abstract).
Dispersal biology of the ophiostomatoid fungi. Taxonomy and Biology of the
Ophiostomatales Symposium, Bad Windsheim, West Germany, August 1990
(with D. Malloch) (Abstract).
Arthropod-mediated dispersal in coprophilous fungi. Fourth International
Mycological Congress, Regensburg, West Germany, August-September 1990 (with
Phylogenetic systematics of ascomycetes. Fungal Holomorph Conference, Newport,
Oregon, August 1992.
Iterative germination in arthropod-associated fungi. MSA, Portland, Oregon,
August 1992 (with D. Malloch) (Abstract).
Morphological convergence among arthropod dispersed fungi studied using cladistic
analysis of molecular data. British Mycological Society-Systematics Association,
Kew, England, November 1992 (with J. W. Spatafora).
Molecular systematics of unitunicate perithecial ascomycetes. First International
Workshop on Ascomycete Systematics. Paris, France, May 1993 (with J. W. Spatafora).
Molecular data sets and broad taxon sampling in detecting morphological
convergence. First International Workshop on Ascomycete Systematics. Paris,
France, May 1993 (with J. W. Spatafora).
Minute mycological mysteries: The influence of arthropods on the lives of
fungi. Presidential Address of the Mycological Society of America. MSA, Athens,
Georgia, June 1993 (Abstract).
Evolution of fungal mutualism with arthropods: involvement of phoresy and
parasitism. Society for Invertebrate Pathology, Ashville, North Carolina,
August 1993 (with J. W. Spatafora).
Phylogenetic reconstruction as an aid to understanding the origins of symbiotic
interactions. Gordon Research Conference on Fungal Metabolism, Hoderness School,
Plymouth, New Hampshire, June 1994.
Phylogeny of filamentous fungi deduced from analysis of molecular characters,
present and future. Fungal Taxonomy and Tropical Mycology: Quo Vadis? 100th
Anniversary Celebration of MUCL, Louvain-la-Neuve, Belgium, June 1994.
Heterochrony in fungal life cycles. IMC5, Vancouver, British Columbia, August
1994 (with K. Jones and J. W. Spatafora) (Abstract).
Supraordinal classification of the ascomycetes inferred from molecular and
other criteria: lichenized and non-lichenized pyrenomycetes. IMC5, Vancouver,
British Columbia, August 1994.
Evolutionary simplification of the life cycles of obligate fungal associates
of arthropods. International Union of Microbiological Societies Congresses,
Mycology Division, Prague, Czech Republic, July 1994 (with K. G. Jones and
S. C. Cassar) (Abstract).
Phylogeny of the mycelial fungi; an evaluation of molecular and morphological
data. VI International Marine Mycology Symposium (a meeting of the British
Mycological Society). Portsmouth, U.K., July 1995 (Abstract).
Key to successful funding--making your research relevant to non-mycologists.
MSA, San Diego, August 1995.
A diversity of ascomycetes associated with so many arthropods. British Mycological
Society, Centenary, Sheffield, April 1996.
NSF Nematode Workshop participant, Tucson, Arizona, July 1997.
Southern Forest Insect Work Conference, Folly Beach, South Carolina, August
Phylogeny of phoretic and parasitic fungi associated with insects. Joint
American Pathological Society/Entomological Society Of America, Las Vegas,
Nevada, November 1998.
Teaching Mycology in China. Beijing, China, Spring 1998 (did not attend).
Fungi that fly: associations of fungi with insects and the consequences.
Plenary Lecture, XVI International Botanical Congress, St. Louis, Missouri,
Food vacuole destiny in a new myxomycete. Texas Society of Electron Microscopists,
Huntsville, Texas, May 1972 (Abstract).
A new species of the genus Licea (Myxomycetes). Alexopoulos Appreciation
Day. Amherst, Massachusetts, June 1972 (Abstract).
Entomogenous fungi associated with termites. Association of Southeastern
Biologists, Blacksburg, Virginia, April 1975 (Abstract).
Use of pigment separation as a taxonomic character in the Trichiaceae
(Myxomycetes). Michigan Academy of Arts and Sciences, Ypsilanti, March
1978 (with A. Busard).
Distribution, host specificity, and variation of Arthrorhynchus nycteribiae
and A. eucampsipodae (Laboulbeniomycetes). Mycological Society
of America, Stillwater, Oklahoma, August 1979 (Abstract).
Myxomycetes in different vegetation zones of Arizona. MSA, Tucson, Arizona,
August 1980 (with R. L. Gilbertson) (Abstract).
A genealogy of American mycologists. MSA, Bloomington, Indiana, August 1981
(with R. L. Gilbertson) (Abstract).
Myxomycete-arthropod associations. BSA(M), University Park, Pennsylvania,
August 1982 (Abstract).
Wood-rotting basidiomycetes associated with Quercus virginiana Lam.
ASB, Lafayette, Louisiana, April 1983 (with R. L. Gilbertson).
Cultural studies of wood-rotting basidiomycetes from the Sonoran Desert.
MSA, Ames, Iowa, June 1983 (with R. L. Gilbertson) (Abstract).
Fungi which decay bark of living live oak trees. MSA, Ames, Iowa, June 1983
(with R. L. Gilbertson) (Abstract).
Distribution and sporulation phenology of myxomycetes in the Sonoran Desert
of Arizona. MSA, Fort Collins, Colorado, August 1984 (with R. L. Gilbertson)
Biogeography of fungal obligate parasites of termites. MSA, Fort Collins,
Colorado, August 1984 (with W. Rossi) (Abstract).
The chemical basis of Melzer's reaction. MSA, Gainesville, Florida, August
1985 (with A. J. Kinney, P. T. Radford, C. M. Dugas, and R. L. Gilbertson)
The exudate of Inonotus dryadeus. MSA, Gainesville, Florida, August
1985 (with G. M. W. Adams) (Abstract).
Pyxidiophora-Thaxteriola. Specialization for mite dispersal. MSA,
Amherst, Massachusetts, August 1986 (with J. R. Bridges, J. C. Moser, and
T. J. Perry) (Abstract).
Relationships of the Laboulbeniales. MSA, Ottawa, Ontario, June 1987
(with D. Malloch) (Abstract).
A reassessment of the genus Pyxidiophora. MSA, Davis, California,
August 1988 (with D. Malloch) (Abstract).
Similarity of the adhesive conidia of Basidiobolus to thalli of Amphoromorpha.
MSA, Toronto, Ontario, August 1989 (with D. Malloch) (Abstract).
The importance of arthropod dispersal in coprophilous habitats. MSA, Toronto,
Ontario, August 1989 (with D. Malloch) (Abstract).
Dispersal biology of Stylopage anomala. MSA, Madison, Wisconsin,
June 1990 (with D. Malloch) (Abstract).
New evidence from an anamorph for the removal of Pyxidiophora from
the Hypocreales, MSA, Madison, Wisconsin, June 1990 (with J. W. Spatafora,
D. Malloch, J. W. Taylor) (Abstract).
Description of a unique cuticular fungus infecting Japanese beetle larvae.
Entomological Society of America, New Orleans, December 1990 (with J. L. Hanula
and T. J. Andreadis) (Abstract).
Nuclear-encoded small-subunit ribosomal DNA and ascomycete evolution. Mycological
Society of America, San Antonio, August 1991 (with J. W. Spatafora) (Abstract).
Convergent evolution among arthropod-dispersed perithecial ascomycetes.
MSA, Portland, Oregon, August 1992 (with J. W. Spatafora) (Abstract).
Taxonomic studies of the form genus Ambrosiella based upon rDNA sequences.
MSA, Portland, Oregon, August 1992 (with S. Cassar and J. W. Spatafora) (Abstract).
Laboulbeniales associated with phoretic mites of dung beetles. MSA,
Portland, Oregon, August 1992 (with K. M. Robertson) (Abstract).
Analysis of Pyxidiophora spp. using rDNA sequences. MSA, Portland,
Oregon, August 1992 (with B. E. Sawyer and J. W. Spatafora) (Abstract).
Monophyly and higher level systematics of ascomycetes. MSA, Portland, Oregon,
August 1992 (with J. W. Spatafora) (Abstract).
Phylogenetic origins of ambrosia fungi in the form genus Ambrosiella.
First International Workshop of Ascomycete Systematics. Paris, France, May
1993 (with S. Cassar).
Life histories of three undescribed species of Pyxidiophora occurring
on beached marine algae. MSA, Athens, Georgia, June 1993 (with D. Malloch)
Assessment of patterns of host-mediated isolation in mutualistic fungi of
ambrosia beetles (Coleoptera; Scolytidae). MSA, Athens, Georgia,
June 1993 (with S. Cassar) (Abstract).
Evolution of conidium types in Pyxidiophora. IMC5, Vancouver, British
Columbia, August 1994 (with H. Johnson, B. Sawyer, K. Jones, and D. Malloch)
Yeast and yeast-like endosymbionts of anobiid beetles. MSA, San Diego, California,
August 1995 (with K. G. Jones and P. F. Dowd) (Abstract).
An investigation of position specificity in Stigmatomyces using molecular
methods. MSA, San Juan, Puerto Rico, June 1998 (with A. Weir) (Abstract).
Molecular phylogeny of Melanospora Corda and similar pyrenomycetous
fungi. XVI International Botanical Congress, St. Louis, Missouri, August 1999
(with N. Zhang) (Abstract).
Yeasts from the gut of Coleoptera: isolation and molecular phylogeny. XVI
International Botanical Congress, St. Louis, Missouri, August 1999 (with S-O
Phylogenetic position of the termite ectoparasitic Termitaria snyderi
and Laboulbeniopsis termitarium. XVI International Botanical Congress,
St. Louis, Missouri, August 1999 (with D. Henk, K. G. Jones, and A. Weir)
Host specificity: Stigmatomyces (Fungi; Ascomycota) parasitic on Ochthera
(Diptera; Ephydridae). XVI International Botanical Congress, St. Louis, Missouri,
August 1999 (with A. Weir and W. Rossi) (Abstract).
Fossil bacteria 3.4 Ga and fungi of unknown age in Archean cherts: distinguishing
microfossils of different ages using light and electron microscopy and TOF-SIMS.
Geological Society of America, Denver, Colorado, October 1999 (with M. M.
Walsh, F. Westfall, A. Steele) (Abstract).
UNIVERSITY DUTIES (selected)
Departmental Seminar Committee, 1976-1977; Biology Faculty Search Committee,
1976-1977; College Cultural Affairs Committee, 1977-1979; Rhodes Scholarship
Selection Committee, 1979-1980; Arts and Sciences Space Committee, 1981-1982;
Departmental Graduate Brochure Committee, 1982, 1987; Departmental Seminar
Chairman, 1982-1983; Departmental Library Liaison, 1982-1990; Basic Sciences
Faculty-Student Affairs Committee, 1983-1984; Basic Sciences Day Committee,
1983-1984; Faculty Senate, 1984-1985; Chairman Evaluation Committee, 1985;
Basic Sciences Faculty Teaching Evaluation Committee, 1985; University Commencement
Committee, 1987; Basic Sciences Teaching Awards Committee, 1987-1988; Agriculture
and Life Sciences Colloquium Committee, 1987-1988; University Honors Board,
1989-1992; Departmental Undergraduate Recruiting Materials Committee, 1989;
Basic Sciences Community Earth Day Committee, 1990; Basic Sciences Tenure
and Promotions Committee, 1991, 1992; Basic Sciences Policy and Planning Committee,
1992-1993; Basic Sciences Tenure and Promotions Committee, Chairman, 1994-1995;
Basic Sciences Committee on Research, 1995-1998; Basic Sciences Safety Committee,
1995-1997; Departmental Chair Search Committee, Chairman, 1995; Systematics
Search Committee, 1997; Systematics Search Committee, Chairman, 1998; Departmental
Newsletter Editor, 1999.